Knowles died in 1933, but this Sunday her life and work will be honoured by a new plaque at the National Botanic Gardens. Better late than never.
She was an expert on lichens, those crusty growths you see on rocks, walls and trees. Significantly, she was the first person to recognise that, at the shore, lichens grow in distinct tidal bands or zones; she discovered this while studying lichens at Howth.
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